1a. |
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1b. |
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2a. |
Stipules 6–20 by 11–37 mm, often long persistent. Petiole upper pulvinus swollen, 5–11 by 3–6 mm. Leaves 14–47 cm long; base cordate; nerves raised beneath, not fading towards the margin |
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2b. |
Stipules 4–8 by 11–18 mm, present in young stages. Petiole upper pulvinus distinct, but not extremely so, 2–6 by 1.5–2.5 mm. Leaves 8–25 cm long; base emarginate to rounded; nerves prominent beneath, fading towards the margin |
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3a. |
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3b. |
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4a. |
Petioles and leaves completely glabrous. Leaf: margin subentire, lowly undulate or lowly glandular-crenate |
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4b. |
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46c. A. octandra var. malesiana |
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5a. |
Leaf: basal glands fading. Glomerules ellipsoid, slightly interruptedly set along the rachis. — Sulawesi: Sula Isl. |
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5b. |
Leaf: basal glands distinct. Glomerules indistinct, continuously set along the rachis. — Philippines |
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6a. |
Branchlets, petiole, and leaf beneath tomentose |
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6b. |
Branchlets, petiole, and leaf beneath glabrous or sparsely puberulous |
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7a. |
Stipules caducous. Leaves glabrous above. Inflorescences 18–47 mm long, 1.5–2.2 mm diam., tomentose; glomerules 1.5–2 by 1–1.5 mm. Stamens 2 or 3, not to slightly exserted, 0.8–1 mm long |
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7b. |
Stipules falcate, often rather persistent. Leaves sparsely tomentose above. Inflorescences 14–59 mm long, 2–3 mm diam., densely (reddish) tomentose; glomerules 2–4 by 1.2–2 mm. Stamens 4, exserted, 1.2–1.8 mm long |
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8a. |
Leaf: dots absent; drying yellowish or greyish green on both sides; young leaves lowly white granular beneath, mature ones smooth |
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8b. |
Leaf: dots distinct, sometimes fading; drying greyish, greenish, or brownish above, brownish beneath; young and mature leaves smooth |
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9a. |
Inflorescences 1–1.5 mm diam.; peduncle 1.5–2.5 mm long, subglabrous; glomerules subglobose, spaced 0.5–2 mm at base |
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9b. |
Inflorescences 2–3.5 mm diam.; peduncle 0.5–1.5 mm long, sparsely puberulous; glomerules ellipsoid, spaced 0.5–1 mm at base |
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10a. |
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10b. |
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11a. |
Branchlets and leaves completely glabrous. Nerves 7–9 pairs. — Moluccas |
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11b. |
Branchlets and leaves on nerves (very) sparsely puberulous beneath. Nerves 9–13 pairs. — Central Sulawesi |
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12a. |
Inflorescences: 28–74 mm long, sparsely puberulous; glomerules distinctly, up to 3 mm, spaced at base. — Philippines |
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12b. |
Inflorescences: 11–26 mm long, subglabrous; glomerules not or slightly, up to 1 mm, spaced at base — Moluccas: Seram |
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13a. |
Leaves 6.5–15 cm long; marginal glands present, but not extremely large; dots densely set |
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13b. |
Leaves 11–20 cm long; marginal glands thick, black, protruding; dots irregularly laxly set |
1a. |
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1b. |
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2a. |
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2b. |
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3a. |
Stipules 6–20 by 11–37 mm, often long persistent. Petiole: upper pulvinus swollen, 5–11 by 3–6 mm. Leaves 14–47 cm long; base cordate; nerves beneath raised, not fading towards the margin |
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3b. |
Stipules 4–8 by 11–18 mm, present in young stages. Petiole: upper pulvinus distinct, 2–6 by 1.5–2.5 mm. Leaves 8–25 cm long; base emarginate to rounded; nerves beneath prominent, fading towards the margin |
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4a. |
Branchlets sparsely puberulous to glabrous. Petiole and leaf glabrous. Leaf: margin subentire, lowly undulate or lowly glandular-crenate. Ovary and fruit sparsely sericeous to glabrous |
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4b. |
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46c. A. octandra var. malesiana |
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5a. |
Leaf: basal glands fading. Ovary and fruit sparsely sericeous (?glabrous). — Moluccas |
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5b. |
Leaf: basal glands distinct. Ovary and fruit glabrous. — Philippines |
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6a. |
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6b. |
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7a. |
Ovary glabrous; stigmas prominent, papillate to laciniate. Young fruits ovoid, 12–16 by 12–16 mm, mature fruits globose, 23–27 by 23–28 mm, punctulate; stigmas prominent, irregularly sculptured and ridged on sutures; pericarp 0.5–1.5 mm thick when immature to 4–6 mm thick when mature |
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7b. |
Ovary glabrescent; stigmas flat, lowly papillate. Young and mature fruits globose, 9–15 by 10–15 mm, smooth, stigmas sunken; pericarp 0.5–2.5 mm thick |
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8a. |
Branchlets, petiole, and leaf beneath tomentose |
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8b. |
Branchlets, petiole, and leaf beneath glabrous or sparsely puberulous |
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9a. |
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9b. |
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10a. |
Ovary ovoid. Fruit ovoid to globose |
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10b. |
Ovary narrowly ovoid. Fruit fusiform. — (Sometimes?/usually?) hermaphrodite |
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11a. |
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11b. |
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12a. |
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12b. |
Branchlets and leaves on nerves (very) sparsely puberulous beneath. Nerves 9–13 pairs. Stigmas 1–1.5 mm long, bifid, not tree-like. Fruits ovoid, 12–17 mm long, slightly beaked. — Central Sulawesi |
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13a. |
Inflorescences 7–16 mm long. Flowers 2- or 3-locular, singly set per bract; petal-like structures absent; stigmas sessile, 0.8–1.2 mm long. — Fruits not beaked, pericarp 0.3–0.5 mm thick |
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13b. |
Inflorescences 24–58 mm long. Flowers 3-locular, sometimes 2 per bract; petal-like structure sometimes distinct; stigmas raised, 2.5–3 mm long. — Fruits unknown |