Species in parentheses and italics key out in this place only as an exception. Species in red are present only in the key, not in the descriptions of the Malesian species (see website on Thai Euphorbiaceae)
SUM = Sumatra, MLY = Malay Peninsula, BOR = Borneo, JAW = Java, PHI = Philippines, SUL = Sulawesi, LSI = Lesser Sunda Islands, MOL = Moluccas, NWG = New Guinea, THA = Thailand.
1a. |
Leaf apex rounded or retuse, more rarely obtuse |
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1b. |
Leaf apex acute, acuminate or caudate |
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Leaves base cordate to rounded, very rarely acute. Sepals free, pubescent outside. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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2b. |
Leaf base acute or obtuse, mostly cuneate. Sepals fused or free, glabrous outside |
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Sepals fused for 2/3 of their length. Leaves obovate to elliptic-oblong, (2–)5–10(–18) by (1–)2.5–4(–8) cm. Leaves: petioles 1–2 mm wide. Fruiting pedicels 1.5–3 mm long. JAW, THA |
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3b. |
Sepals free or nearly so. Leaves spathulate, (1.5–)2–5(–7) by (0.7–)1–2.5(–3.5) cm. Leaves: petioles less than 1 mm wide; fruiting pedicels 0.2–1 mm long. MOL, NWG |
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Midvein in dry material sharply raised adaxially (Fig. 2E), distinctly perceptible to the touch. Inflorescences simple |
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4b. |
Midvein in dry material impressed to slightly raised adaxially (Fig. 2C–D), not distinctly perceptible to the touch. Inflorescences simple or branched |
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Leaves 6–7.5 cm long. Ovaries glabrous. Fruits 5–6(–8) by 4–5(–6) mm. Infructescences 4–8cm long. THA |
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5b. |
Leaves 13–45 cm long. Ovaries pubescent. Fruits 8–16 by 5–9 mm. Infructescences 7–50 cm long |
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Young twigs, Leaves: petioles and leaves hairy, glabrescent. Stipules usually foliaceous, more than 2 mm wide. Leaves 15–45 by 5–18 cm. Sepals fused to halfway; disc glabrous to pilose at the margin. Inflorescences 15–20 cm long, infructescences 25–50 cm long. Fruits glabrous or with stellate hairs, 12–16 mm long, style subterminal. MLY |
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6b. |
Young twigs, Leaves: petioles and leaves glabrous. Stipules linear, up to 1.5 mm wide, caducous. Leaves 13–27 by 3.5–8 cm. Sepals fused only at the base; disc glabrous. Inflorescences c. 3 cm, infructescences up to 15 cm long. Fruits pilose, with simple hairs, 8–11 mm long, style terminal. MLY,THA |
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Inflorescences at least partly cauline |
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7b. |
Inflorescences axillary (sometimes aggregated at the end of the branch) |
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Petioles (at least of some leaves) more than 10 mm long, no wider than 2 mm in the middle, basally and distally usually slightly pulvinate and sometimes geniculate |
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8b. |
Petioles shorter than 10 mm (if longer, then more than 2 mm wide), basally and distally not pulvinate or geniculate |
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Disc hairy. All leaf veins, including the very fine veinlets, in dry material clearly visible abaxially. Leaves drying yellowish brown to yellowish green. Pubescence of inflorescence whitish. Inflorescences simple, fascicled. SUM, MLY, BOR, PHI, THA |
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9b. |
Disc glabrous (sometimes with hairs originating from the base of the disc). Only the coarse leaf veins in dry material visible abaxially. Leaves drying grey to reddish brown. Pubescence of inflorescence ferrugineous. Inflorescences simple or branched. SUM, MLY, BOR |
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Tertiary veins close together, more than 10 between every two secondary veins (Fig. 2Aa). Infructescences 8–25(–69) cm long, up to 13 per fascicle. Leaves: petioles 1.5–4 mm wide. Leaves (7–)13–25(–50) by (2.5–)4–8(–15) cm. Stipules 8–20 by 1–4 mm |
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10b. |
Tertiary veins widely spaced, less than 9 between every two secondary veins (Fig. 2Ab). Infructescences 4–15 cm long, up to 7 per fascicle. Leaves: petioles up to 2 mm wide. Leaves (5–)8–14(–19) by (2–)3–5(–9.5) cm. Stipules 2–10 by 0.5–1.5 mm |
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Fruits 4–8(–10) mm long, lenticular to ellipsoid, rarely ovate and beaked, usually laterally compressed, areolate. Stigmas 4–6. Ovaries hairy to nearly glabrous. Infructescences 10–25(–69) cm long, flexible. Indumentum of leaves sparse abaxially, indumentum of inflorescence axes and calyx moderately dense. SUM, MLY, BOR, PHI, THA |
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11b. |
Fruits 10–14 mm long, elongate-ellipsoid, more or less tetragonal, not or hardly compressed, reticulate. Stigmas 6–16. Ovaries densely hairy. Infructescences 8–13 cm long, robust. Indumentum of leaves usually dense abaxially, indumentum of inflorescence axes and calyx extremely dense. BOR |
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Abaxial leaf surfaces hairy all over |
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12b. |
Abaxial leaf surfaces glabrous |
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Indumentum ferrugineous. Fruits 4–5 by c. 3 mm. Style terminal, usually wide. Inflorescences 2–4 cm long, infructescences 3–8 cm long; disc glabrous (sometimes with hairs originating from the base of the disc). MLY, THA |
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13b. |
Indumentum whitish to ochraceous. Fruits 6–8 by 4–6 mm. Style lateral or subterminal, very distinct, thin. Inflorescences (4–)7–8 cm long, infructescences 8–23 cm long; disc glabrous, or glabrous only at the sides, hairy on top. LSI, MOL, SUL |
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Fruits 6–8 by 4–6 mm, style subterminal to lateral, rarely terminal. Sepals free or nearly so; disc hairy all over. Stigmas (2–)3–4, long (up to 1.5 mm) and conspicuous. SUM, MLY, THA |
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14b. |
Fruits 7–10 by 5–7 mm, style terminal to slightly subterminal. Sepals fused for 2/3–3/4 of their length; disc glabrous (sometimes with hairs originating from the base of the disc). Stigmas 5–8(–10), not conspicuously long. PHI |
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Stipules foliaceous, more than 4 mm wide |
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15b. |
Stipules not foliaceous, up to 4 mm wide |
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Leaves: petioles (at least of some leaves) more than 10 mm long, no more than 2 mm wide in the middle, basally and apically slightly pulvinate and geniculate |
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16b. |
Leaves: petioles up to 10 mm long (if longer, more than 2 mm wide), not pulvinate or geniculate |
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Stipules spathulate, cordate or kidney-shaped, apex rounded to acute, base symmetrical. Leaves (6–)10–14(–22) cm long, up to 3 times as long as wide, chartaceous to membranaceous; bracts c. 0.5 mm long. Sepals fused for 2/3 of their length. Inflorescences usually consisting of 5–8 branches, rarely simple. SUM, JAW, LSI |
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17b. |
Stipules ovate, apex caudate, base asymmetrical. Leaves 13–21 cm long, 3–4 times as long as wide, subcoriaceous to chartaceous; bracts 0.7–1 mm long. Sepals fused only at the base. Inflorescences simple or once-branched. MLY |
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Indumentum, especially of young parts and inflorescences, ferrugineous. Leaves drying reddish brown |
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18b. |
Indumentum whitish or absent. Leaves drying olive-green to reddish brown |
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Sepals fused for 2/3–3/4 of their length; disc ferrugineous-pubescent. Fruits terete or hardly compressed. Mostly rheophytic. BOR, PHI (Palawan), SUL |
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19b. |
Sepals free or nearly so; disc glabrous to ferrugineous-pubescent. Fruits laterally or dorsiventrally compressed, or terete |
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Stipules subulate to falcate, rarely elliptic with an asymmetrical base, strongly parallel-veined, coriaceous, no wider than 10 mm, at least twice as long as wide. Leaves: petioles 2–5 mm wide. Leaves (8–)15–30(–60) cm long. Tertiary veins percurrent, close together (Fig. 2Aa). Inflorescences 3–30 cm long; flowers set densely, touching each other. Infructescences 7–65 cm long. Ovaries ovoid to falcate, densely appressed-pubescent; disc glabrous or hairy. Fruits laterally compressed. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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20b. |
Stipules cordate to elliptic, base more or less symmetrical, reticulately veined, chartaceous, up to 18 mm wide, hardly longer to several times as long as wide. Leaves: petioles c. 1 mm wide. Leaves (5–)6–14(–20) cm long. Tertiary veins weakly percurrent (Fig. 2Ab) to reticulate (Fig. 2Ac). Inflorescences 1.5–6 cm long, lax; flowers widely spaced, not touching each other. Infructescences 4–16 cm long. Ovaries globose, glabrous, rarely pilose; disc glabrous. Fruits laterally or dorsiventrally compressed |
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Leaves more than 2 cm wide, less than 5 times as long as wide. SUM, MLY,BOR, THA |
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21b. |
Leaves less than 2 cm wide, more than 5 times as long as wide. Rheophyte. BOR |
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Fruits terete, (3–)4–6(–8) mm long. Stipules, Leaves: petioles and abaxial leaf surfaces usually hairy. Sepals often glandular-fimbriate. Inflorescences simple to consisting of up to 13 branches. THA |
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22b. |
Fruits laterally compressed, 6–18 mm long. Stipules, Leaves: petioles and abaxial leaf surfaces usually glabrous. Sepals never glandular-fimbriate. Inflorescences simple or consisting of up to 3 branches |
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Fruits 6–8 by 4–6 mm. Inflorescences 4–10 cm long, infructescences 4–15 cm long; fruiting pedicels 0.5–2.5 mm long. Leaves drying greyish green to reddish brown, (5–)9–14(–19) cm long, 1.9–4.3 times as long as wide. Young twigs light grey. Stipules up to 5 mm wide. Disc and ovaries always hairy. SUM, MLY, THA |
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23b. |
Fruits 8–18 by 6–11 mm. Inflorescences 10–30 cm long, infructescences 10–50 cm long; fruiting pedicels 1–14 mm long. Leaves drying olive-green, (9–)20–30(–40) cm long, 3.2–9.2 times as long as wide. Young twigs brown. Stipules up to 30 mm wide; disc and ovaries usually glabrous. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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Bracts more than 1.5 mm long, conspicuous even in very young inflorescences, less so in old infructescences. Leaves abaxially hairy |
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24b. |
Bracts up to 1.5 mm long. Leaves abaxially hairy or glabrous |
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Indumentum, especially of young parts, densely hirsute, whitish to ochraceous, never ferrugineous. Leaves drying olive-green. Fruits lenticular, obliquely ellipsoid or bean-shaped, 4–7 by 4–6 mm, with a distinctly asymmetrical base and lateral styles, laterally compressed. Sepals 5–8. Infructescences 6–15 cm long. Most leaves 12–18 by 4–6 cm, tertiary veins percurrent, close together (Fig. 2Aa); disc glabrous. SUM, MLY, BOR (Anambas & Natuna Isl.), JAW, THA |
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25b. |
Indumentum, especially of young parts, ferrugineous. Leaves drying reddish brown. Fruits ellipsoid or obliquely ovate (mango-shaped) (7–)9–10(–20) by 5–9 mm, with a symmetrical or asymmetrical base and terminal to lateral styles, not, laterally or dorsiventrally compressed. Sepals 4–6. Infructescences 10–65 cm long. Most leaves 15–30 by 5–12 cm, tertiary veins reticulate (Fig. 2Ac) to percurrent (Fig. 2Aa), close together or widely spaced; disc hairy or glabrous |
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Indumentum of all parts rather long and shaggy; sometimes, especially on fruits, with some stellate hairs. Fruits ellipsoid, not or laterally or dorsiventrally compressed. Tertiary veins reticulate (Fig. 2Ac) to weakly percurrent (Fig. 2Ab), widely spaced; disc hairy at the margin. Usually above 1000 m altitude. MLY |
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26b. |
Indumentum of all parts rather short and velvety; without stellate hairs. Fruits obliquely ovate (mango-shaped), always laterally compressed. Tertiary veins percurrent, close together (Fig. 2Aa); disc hairy or glabrous. Indifferent to altitude. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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Leaves: petioles (at least of some leaves) more than 10 mm long, no wider than 3 mm in the middle, basally and apically usually slightly pulvinate and sometimes geniculate |
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27b. |
Leaves: petioles up to 10 mm long (if longer, more than 3 mm wide, and leaf blades more than 25 cm long, basally and distally never pulvinate or geniculate) |
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Abaxial leaf surfaces hairy all over |
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28b. |
Abaxial leaf surfaces glabrous, or hairy along the major veins only |
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Disc glabrous (sometimes with hairs originating from the base of the disc). Fruits less than 3 mm or more than 7 mm long, glabrous |
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29b. |
Disc hairy all over or at least at the margin. Fruits 3–6 mm long, pilose or glabrous |
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Fruits 1.5–2.5 mm long, style distinctly lateral. Inflorescences branched. Leaf indumentum abaxially not distinctly velvety to the touch. Leaves: petioles 5–13 by 0.8–1 mm. Most leaves 10–13 by 3.5–5 cm. BOR |
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30b. |
Fruits 8–10 mm long, style terminal or subterminal. Inflorescences simple. Leaf indumentum abaxially distinctly velvety to the touch. Leaves: petioles 12–35 by 1.2–2 mm. Most leaves 16–21 by 6–8 cm. MLY |
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Fruits with lateral style and asymmetrical base, bean-shaped to obliquely lenticular. Leaves: petioles 1.5–3 mm wide. Stipules 1–2 mm wide. Leaves (8–)13–17(–23) by (3–)5–8(–12) cm. PHI |
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31b. |
Fruits with terminal style and symmetrical base, lenticular or ellipsoid. Leaves: petioles 0.5–1.5 mm wide. Stipules 0.5–1 mm wide. Leaves (2–)4–11(–16) by (2–)3– 5(–9) cm |
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Sepals fused to halfway. Ovaries (and to a lesser extent also the fruit) elongate-ellipsoid. Inflorescences simple. Leaves (1.3–)1.8–2.4(–4) times as long as wide. PHI, LSI |
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32b. |
Sepals free. Ovaries ovoid to globose, fruits lenticular, more rarely ellipsoid. Inflorescences consisting of 3–20 branches. Leaves (1–)1.3–1.7(–2.25) times as long as wide. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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Fruits with a distinctly lateral style (Fig. 3L); fruiting pedicels 0.5–5 mm long |
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33b. |
Fruits with a terminal (Fig. 3G) to subterminal (Fig. 3J) style; fruiting pedicels 0–2mm long |
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Fruits 1.5–2.5 by 1–2 mm. BOR |
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34b. |
Fruits 3–15 by 2–12 mm |
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Sepals free. SUM, MLY, BOR |
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35b. |
Sepals fused for at least half their length |
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Disc glabrous (sometimes with hairs originating from the base of the disc; Fig. 3D) |
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36b. |
Disc hairy all over (Fig. 3F), or at least at the margin (Fig. 3E) |
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Leaves ovate to elliptic, base acute to obtuse. Leaves: petioles 0.5–1 mm wide. Young twigs first dark brown, then light grey to whitish. Inflorescences 2–5 cm long, simple, solitary or 2 per fascicle. Fruits 5–8 by 4–6 mm. Ovaries glabrous to sparsely pilose. SUM, MLY, THA |
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37b. |
Leaves usually oblong, base rounded to truncate, rarely obtuse. Leaves: petioles 0.7–2 mm wide. Young twigs brown. Inflorescences 5–10 cm long, usually consisting of 5–8 very regular branches, solitary. Fruits 3–5 by 2–3 mm (unknown for Antidesma cuspidatum var. orthocalyx) |
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Inflorescences 5–6 cm long. Ovaries glabrous. Stipules (2–)5–20 mm wide. Fruits 3–5 by 2–3 mm. SUM, JAW, LSI |
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38b. |
Inflorescences 8–10 cm long. Ovaries pilose. Stipules and fruits unknown. SUM |
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Exocarp in dry material loosely enfolding the endocarp, thick, neither areolate nor reticulate, without ridges or keels, brittle, minutely scaly (Fig. 5C). Fruits globose, not or hardly compressed. Infructescences consisting of 1–4 branches, axes robust and rigid. MOL, NWG |
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39b. |
Exocarp in dry material closely enfolding the endocarp, thin, reticulate, areolate or wrinkled (fleshy when fresh), usually keeled at the suture, not brittle or scaly (Fig. 5D). Fruits laterally compressed. Infructescences consisting of up to 10 branches, axes less robust and rigid |
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Leaves: petioles 0.8–1.2(–2) mm wide. Leaves chartaceous to membranaceous, without domatia, base obtuse to truncate, rarely acute. Young twigs often very light brown to whitish. Fruits 3–4 mm across; ovaries and fruits hairy.PHI |
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40b. |
Leaves: petioles 1–2(–3) mm wide. Leaves coriaceous to subcoriaceous, sometimes with domatia, base acute to rounded. Young twigs brown. Fruit 3–9 mm across; ovaries and fruits glabrous to sparsely pilose |
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Bracts 0.4–0.8 by 0.3–0.6 mm. Infructescences 4–17 cm long, consisting of 2–10 branches, rarely simple. Domatia in the axils between the midvein and secondary veins abaxially (Fig. 2B) usually present. Occasionally cultivated. BOR, PHI, SUL, MOL, NWG |
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41b. |
Bracts 1–1.5 by 0.7–1 mm. Infructescences 4 cm long, simple. Domatia absent. PHI |
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Sepals free, pubescent or hispid outside. Indumentum of inflorescence axes usually ferrugineous |
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42b. |
Sepals fused for 1/2 of their length or more (fruiting calyx splits irregularly in Antidesma curranii), glabrous to pilose outside. Indumentum of inflorescence axes usually whitish |
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Ovaries and fruits glabrous. Fruits (3–)4–8(–13) by 4–6 mm. Leaves: petioles 1–2 mm wide. Stigmas 4–10. Infructescences consisting of 1–10 branches. Leaves (6–)10–15(–21) cm long, coriaceous, drying grey to reddish brown adaxially, reddish brown abaxially. SUM, MLY, BOR |
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43b. |
Ovaries and fruits hairy. Fruits 3–4(–5) by 2.5–3(–3.5) mm. Leaves: petioles 0.7–1 mm wide. Stigmas 2–4. Infructescences consisting of (1–)10–20 branches. Leaves (2–)4–6(–16) cm long, chartaceous to coriaceous, drying olive-green. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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Fruits 4–5 by 3–4 mm; calyx in flower fused to halfway, splitting irregularly in fruit, 1–1.5 mm long, much longer than the disc. Inflorescences simple or consisting of up to 6 branches, solitary. PH |
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44b. |
Fruits 2–4 by 1–3 mm; calyx in flower fused for (1/2–)2/3–4/5 of its length, not split in fruit, 0.5–1 mm long, extending to about the same length as the disc. Inflorescences consisting of 2–20 branches, often 2–3per fascicle |
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Calyx bowl-shaped, 0.5–0.7 by c. 1 mm. Fruits distinctly laterally compressed, 3–4 by c. 3 mm; fruiting pedicels 1–2 mm long. Leaves membranaceous to subchartaceous, usually with domatia in the axils between the midvein and secondary veins abaxially (Fig. 2B), base acute to obtuse, more rarely rounded. Young twigs brown. SUL, LSI, MOL |
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45b. |
Calyx urceolate to cupular, 0.5–1 by 0.7–1 mm. Fruits not or slightly compressed, 2–3(–4) by 1–2.5 mm; fruiting pedicels 0.2–2 mm long. Leaves chartaceous to subcoriaceous, without domatia, base often obtuse to rounded. Young twigs light brown to whitish |
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Disc ferrugineous-pubescent, hairs as long as to slightly longer than the disc itself. Fruits with terminal style, 2–3(–4) by 1–2 mm; fruiting pedicels 0.2–0.5(–1.5) mm long. PHI |
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46b. |
Disc sparsely whitish-pilose, hairs much shorter than the disc. Fruits with subterminal style, 3.5 by 2–2.5 mm; fruiting pedicels 1–2 mm long. PHI |
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Disc glabrous (sometimes with hairs originating from the base of the disc; Fig. 3D) |
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47b. |
Disc hairy all over (Fig. 3F), or at least at the margin (Fig. 3E) |
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Fruits 1.5–2.5 mm long, glabrous, style lateral. Sepals fused for c. 2/3 of their length. BOR |
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48b. |
Fruits more than 3 mm long, glabrous or hairy, style terminal to lateral. Sepals free or fused |
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Sepals fused for more than half of their length |
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49b. |
Sepals free to fused for up to half of their length (if fused to the middle, then both leads are correct) |
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Fruits roughly bean-shaped, with a lateral style and an asymmetrical base, 5–8 by 4–6 mm. Leaves chartaceous to submembranaceous. SUM, MLY,THA |
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50b. |
Fruits ellipsoid, with a terminal to slightly subterminal (sometimes lateral in Antidesma stipulare) style and a symmetrical base, 3–18 by 4–11 mm. Leaves chartaceous to coriaceous |
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Fruits (3–)4–6(–8) by 3–4(–6) mm, laterally compressed or terete. Leaves with or without domatia, pilose at least along the major veins |
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51b. |
Fruits 5–18 by 4–11 mm, laterally compressed. Leaves without domatia, completely glabrous or pubescent |
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Inflorescences 2–3 cm long, infructescences 2–5(–8) cm long. Leaves with domatia between the midvein and secondary veins abaxially. Calyx urceolate, lobed for up to 1/3 of its length. JAW, THA |
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52b. |
Inflorescences 3–10 cm long, infructescences 6–20 cm long. Leaf domatia present or absent; calyx urceolate or cupular, lobed for 1/4 to almost all of its length |
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Calyx urceolate, lobed for 1/4–1/2 of its length, margin never glandular-fimbriate. Leaves often with domatia, ochraceous-pilose at least along the veins abaxially, higher venation finely tessellated (Fig. 9D). Fruits without or with inconspicuous white pustules when dry. THA |
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53b. |
Calyx cupular, lobed for c. 1/2 of its length, margin often glandular-fimbriate. Leaves without domatia, usually glabrous except along the major veins, higher venation not conspicuously tessellated (Fig. 9A). Fruits usually with conspicuous white pustules when dry |
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Plant rheophytic, leaves 4–10 times as long as wide. SUM, MLY, BOR, THA |
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54b. |
Plant not rheophytic, leaves (2–)2.5–3.5(–6.6) times as long as wide. SUM, MLY,BOR, JAW,PHI, SUL, LSI, MOL, THA |
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Midvein sharply keeled abaxially (Fig. 2F). Fruits 10–15 by 7–11 mm, thinly appressed-puberulent. Leaves coriaceous. BOR |
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55b. |
Midvein rounded abaxially (Fig. 2C–E). Fruits 5–18 by 4–11 mm, glabrous or hairy.Leaves coriaceous |
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Leaves drying dark reddish brown or purplish grey to greyish brown, usually elliptic. Young twigs and old Leaves: petioles whitish. Stigmas 5–10. Inflorescences 2–4 cm long, infructescences 3–14 cm long. Fruiting pedicels 0–1.5 mm long. PHI |
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56b. |
Leaves drying olive-green to yellowish brown, usually oblong. Young twigs brown. Stigmas 3–5(–6). Inflorescences (4–)8–35 cm long, infructescences 10–75 cm long; fruiting pedicels 0–14 mm long |
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Sepals fused for 2/3–3/4 of their length; calyx in flower and fruit cupular, 1–1.5 mm long, with long ferrugineous hairs at the base inside. Fruits 5–11 by 4–7 mm. Stipules early-caducous, 4–6 mm long. Leaves (5–)10–18(–32) cm long |
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57b. |
Sepals fused for c. 1/2 of their length; calyx in flower and fruit bowl-shaped to cupular, up to 1 mm long, indumentum short or absent, whitish to ochraceous. Fruits 8–18 by 6–11 mm. Stipules usually persistent, 5–60 mm long. Leaves (9–)20–30(–60) cm long |
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Abaxial leaf surfaces glabrous except sometimes along the midvein. Ovaries glabrous. Often cultivated. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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58b. |
Abaxial leaf surfaces pubescent all over. Ovaries sparsely pilose. THA |
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Fruiting pedicels 0–1(–3) mm long. Fruits ovate to ellipsoid or lenticular, often apiculate, with a terminal to subterminal style. Ovaries hairy. Leaves usually drying yellowish brown. SUM, MLY, BOR, THA |
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59b. |
Fruiting pedicels 1–14 mm long. Fruits obliquely ovate (mango-shaped), with a lateral to subterminal style. Ovaries glabrous, very rarely hairy. Leaves drying olive-green. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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Indumentum, especially of young parts and inflorescences, ferrugineous to ochraceous |
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60b. |
Indumentum whitish or absent |
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Fruits 4–5 by 3 mm; fruiting pedicels c. 1 mm long; calyx urceolate, sepals fused for c. 1/2 of their length. Ovaries almost cylindrical. Leaves ferrugineous-pubescent all over abaxially, distinctly velvety to the touch. Leaves up to 18 cm long. MLY, THA |
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61b. |
Fruits 5–20 by 3–9 mm; fruiting pedicels 1–23 mm long; calyx cupular, sepals free to fused for c. 1/2 of their length, calyx not urceolate. Ovaries falcate, ovoid or globose. Leaves abaxially pubescent to glabrous. Leaves up to 60 cm long |
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Fruits obliquely ovate (mango-shaped) to falcate (sometimes elongate-ellipsoid in rheophytes), always laterally compressed, hairy (old fruits sometimes almost glabrous). Ovaries ovoid to falcate, densely appressed-pubescent. Inflorescences 3–30 cm long, infructescences 7–65 cm long. Leaves: petioles (1–)2–5 mm wide. Leaves (8–)15–30(–60) cm long. Tertiary veins percurrent, close together (Fig. 2Aa) |
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62b. |
Fruits ellipsoid, globose, lenticular, ovate or obovate, compressed or terete, glabrous or hairy. Ovaries globose to lenticular, glabrous or spreading-hirsute. Inflorescences 1.5–8 cm long, infructescences 3–24 cm long. Leaves: petioles 1–2 mm wide. Leaves (3–)6–14(–20) cm long. Tertiary veins weakly percurrent (Fig. 2Ab) to reticulate (Fig. 2Ac), widely spaced |
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Plant rheophytic, leaves more than 6 times as long as wide. Leaves: petioles 1–3 mm wide. Inflorescences 3–5(–20) cm long, infructescences 7–20 cm long. Fruits 7–9 by 3–6 mm. BOR |
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63b. |
Plant not rheophytic, leaves less than 6 times as long as wide. Leaves: petioles 2–5 mm wide. Inflorescences (5–)10–30 cm long, infructescences 10–30 (–65) cm long. Fruits (7–)9–20 by 5–9 mm. SUM, MLY, BOR, JAW,PHI, SUL, LSI, MOL, THA |
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Leaves: petioles 1–2 mm wide. Leaves membranaceous to chartaceous, spreading-hirsute all over abaxially, drying olive-green to reddish green. Fruits ellipsoid to lenticular, laterally compressed, usually conspicuously beaked, pilose, rarely nearly glabrous, usually conspicuously and coarsely white-pustulate. Inflorescences (4–)7–8 cm long. Ovaries spreading-hirsute; calyx 1 by 1.5–2 mm. LSI, MOL, SUL |
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64b. |
Leaves: petioles c. 1 mm wide. Leaves chartaceous to coriaceous, glabrous or pilose only along the major veins (rarely all over) abaxially, drying dark reddish brown. Fruits ellipsoid, globose, ovoid or obovoid, laterally or dorsiventrally compressed, or terete, not beaked, glabrous, rarely thinly puberulent, rarely white-pustulate. Inflorescences 1.5–6 cm long. Ovaries glabrous, rarely pilose; calyx 0.3–0.8 by 0.8 mm |
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Stipules 0.5–1mm wide. SUM, MLY,BOR |
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65b. |
Stipules 1.5–18 mm wide |
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Plant rheophytic, leaves more than 5 times as long as wide. Infructescences 3–5 cm long. BOR |
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66b. |
Plant not rheophytic, leaves less than 5 times as long as wide. Infructescences 6–9 cm long. SUM, MLY, BOR, THA |
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Young twigs light brown to whitish. Leaves drying reddish brown to grey, often with conspicuous dense intersecondary and perpendicular tertiary veins near the midvein |
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67b. |
Young twigs brown. Leaves drying olive-green, without conspicuous perpendicular intersecondary and tertiary veins near the midvein |
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Fruiting pedicels 0–1 mm long. Higher order veins in dry material distinctly prominent on both leaf surfaces. Stipules caducous, subulate. Leaves drying reddish brown. Fruits ellipsoid, distinctly laterally compressed, with a subterminal to terminal style. Usually at 400–1400 m altitude. SUM, JAW |
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68b. |
Fruiting pedicels 1–18 mm long. Higher order veins in dry material much less prominent adaxially. Stipules usually persistent, elliptic, linear or subulate, often falcate. Leaves drying reddish brown to greyish. Fruits ellipsoid, globose, ovate or obovate, dorsiventrally or laterally compressed, or terete, with a subterminal to lateral style. Indifferent to altitude. SUM, MLY, BOR |
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Fruits 8–18 by 6–11 mm, reticulate, laterally compressed. Sepals never glandular-fimbriate. Flowers 4–5-merous. Inflorescences simple, 8–35 cm long, infructescences 10–75 cm long. Stipules 5–60 by 0.5–30 mm. Leaves (9–)20–50 by (1.5–)5–9(–14) cm, midvein flat to slightly raised, rarely slightly impressed adaxially. Stipules, Leaves: petioles and inflorescence axes usually glabrous |
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69b. |
Fruits 3–8 by 2.5–6 mm, areolate, not or laterally, or slightly dorsiventrally compressed. Sepals often glandular-fimbriate. Flowers 3–5(–7)-merous. Inflorescences simple or branched, 2–10 cm long, infructescences 2.5–20 cm long. Stipules 1.5–7(–13) by 0.5–1.5(–3) mm. Leaves 1.5–15(–30) by 0.4–6(–12) cm, midvein impressed adaxially. Stipules, Leaves: petioles and inflorescence axes usually hairy |
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Fruiting pedicels 0–1(–3) mm long. Fruits ovate to ellipsoid or lenticular, often apiculate, with a terminal to subterminal style. Ovaries hairy. Leaves usually drying yellowish brown. SUM, MLY, BOR, THA |
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70b. |
Fruiting pedicels 1–14 mm long. Fruits obliquely ovate (mango-shaped), with a lateral to subterminal style. Ovaries glabrous, very rarely hairy. Leaves drying olive-green. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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Plant rheophytic, leaves 4–10 times as long as wide or narrower than 2 cm. Fruits terete |
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71b. |
Plant not rheophytic, leaves (1.2–)2.5–3.5(–6.6) times as long as wide. Fruits compressed or terete |
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Most leaves 5–11 by 1–2 cm. Leaf apex acuminate. SUM, MLY, BOR, THA |
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72b. |
Most leaves 3–6 by 0.4–1.2 cm. Leaf apex acute, sometimes with a rounded apiculum. THA |
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Fruits not or hardly laterally compressed, with a terminal style |
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73b. |
Fruits laterally compressed, with a lateral to terminal style |
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Calyx urceolate, lobed for 1/4–1/2 of its length, margin never glandular-fimbriate. Leaves often with domatia, ochraceous-pilose at least along the veins abaxially, higher venation finely tessellated (Fig. 9D). Fruits without or with inconspicuous white pustules when dry. THA |
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74b. |
Calyx cupular, lobed for about half of its length, margin often glandular-fimbriate. Leaves without domatia, usually glabrous except along the major veins, higher venation not conspicuously tessellated (Fig. 9A). Fruits usually with conspicuous white pustules when dry. SUM, MLY, BOR, JAW,PHI, SUL, LSI, MOL, THA |
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Abaxial leaf surfaces spreading-hirsute all over; calyx hirsute outside. Fruits pilose to nearly glabrous, with a lateral style, usually with conspicuous white pustules when dry. Infructescences (4–)8–24 cm long. LSI, MOL, SUL |
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75b. |
Abaxial leaf surfaces glabrous, or pilose only along the major veins. Calyx glabrous outside. Fruits glabrous, with a subterminal to terminal style and only occasionally with rather fine white pustules when dry.Infructescences 4–9cm long |
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Leaves chartaceous to subcoriaceous, often conspicuously shiny, narrowly elliptic-oblong to ovate or obovate, most leaves 6–10 by 2–3.5 cm. Leaves: petioles 0.5–1 mm wide. Leaves: petioles, inflorescences and pedicels slender; fruiting pedicels (2–)3–6 mm long. Sepals free. MLY,PHI, THA |
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76b. |
Leaves membranaceous to chartaceous, not conspicuously shiny, oblong, more rarely elliptic or obovate, most leaves 10–15 by 3–5 cm. Leaves: petioles 1–2 mm wide. Leaves: petioles, inflorescences and pedicels not particularly slender; fruiting pedicels 1–3 mm long. Sepals free to fused for up to half of their length. JAW,LSI, SUL |
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Sepals fused for more than half of their length |
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77b. |
Sepals free to fused for up to half of their length |
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Infructescences up to 10 cm long, simple or branched |
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78b. |
Infructescences more than 10 cm long, always simple |
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Style in flower and fruit distinctly lateral (Fig. 3L) |
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79b. |
Style in flower and fruit terminal (Fig. 3G) to subterminal (Fig. 3J) |
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Fruits beaked, longer than wide (including the beak). Leaves membranaceous to chartaceous. Midvein usually slightly raised, rarely flat, never impressed adaxially. Infructescences 7–10 cm long. SUM, MLY, THA |
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80b. |
Fruits not beaked, as long as wide or wider than long. Leaves chartaceous to coriaceous. Midvein impressed, flat or rarely slightly raised adaxially. Infructescences 2–7(–10) cm long |
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Exocarp in dry material loosely enfolding the endocarp, thick, neither areolate nor reticulate, without ridges or keels, brittle, minutely scaly. Fruits 5–12 mm across, globose, not compressed. MOL, NWG |
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81b. |
Exocarp in dry material closely enfolding the endocarp, thin, reticulate, areolate or wrinkled (fleshy when fresh), not brittle or scaly. Fruits 3–9 mm across, laterally compressed |
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Bracts 0.4–0.8 by 0.3–0.6 mm. Infructescences 4–17 cm long, consisting of 2–10 branches, rarely simple. Domatia in the axils between the midvein and secondary veins abaxially (Fig. 2B) usually present. Occasionally cultivated. BOR, PHI, SUL, MOL, NWG |
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82b. |
Bracts 1–1.5 by 0.7–1 mm. Infructescences c. 4 cm long, simple. Domatia absent. PHI |
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Fruits 10–11 mm long, beaked. Leaves membranaceous to chartaceous. Midvein usually slightly raised, rarely flat, never impressed adaxially. Infructescences 7–10 cm long. SUM, MLY, THA |
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83b. |
Fruits up to 8 mm long, not beaked. Leaves chartaceous to subcoriaceous. Midvein impressed to flat adaxially. Infructescences 2–9 cm long |
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Fruits pilose; calyx pubescent outside. Leaf base cordate to rounded. Sepals 5 0r 6. PHI, LSI |
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84b. |
Fruits glabrous; calyx glabrous outside. Leaf base acute to obtuse, rarely rounded. Sepals 4 or 5 |
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Calyx urceolate, lobed for no more than 1/3 of its length. Leaves sometimes with domatia in the axils between the midvein and secondary veins abaxially (Fig. 2B) |
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85b. |
Calyx cupular, usually lobed for more than 1/3 of its length. Leaf domatia absent |
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Fruiting pedicels 1.5–3 mm long. Leaves obovate to elliptic-oblong, margins basally straight to convex, abaxial leaf surfaces pilose to pubescent at least around the domatia, very rarely completely glabrous. Stipules usually persistent, 3–7 by 1–2 mm. Ovaries and fruits laterally compressed or not. JAW, THA |
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86b. |
Fruiting pedicels 0.5–1.3 mm long. Leaves elliptic to ovate, margins often basally concave, abaxial leaf surfaces glabrous except for sometimes some short hairs along the major veins. Stipules early-caducous, c. 0.5 by 0.5 mm. Ovaries and fruits laterally compressed. PHI |
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Fruits laterally compressed. Infructescences 3–8 cm long; disc indumentum ferrugineous; fruiting pedicels 0–2 mm long |
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87b. |
Fruits terete. Infructescences 6–20 cm long; disc indumentum whitish. Fruiting pedicels 1–4 mm long |
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Leaves: petioles 3–15 mm long. Leaves drying olive-green; calyx in fruit 1–1.5 by 1.5–2 mm; fruiting pedicels 0.5–2 mm long. PHI |
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88b. |
Leaves: petioles 1–3 mm long. Leaves drying reddish brown; calyx in fruit c. 0.4 by 1 mm; fruiting pedicels 0–0.4 mm long. SUL |
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Plant rheophytic, leaves 4–10 times as long as wide. SUM, MLY, BOR, THA |
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89b. |
Plant not rheophytic, leaves (2–)2.5–3.5(–6.6) times as long as wide. SUM, MLY,BOR, JAW,PHI, SUL, LSI, MOL, THA |
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Fruits 8–18 by 6–11 mm, always laterally compressed. Indumentum absent or whitish, never ferrugineous. Leaves drying olive-green to yellowish brown. Young twigs brown |
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90b. |
Fruits 3–8 by 2.5–7 mm, not or hardly compressed. Indumentum ferrugineous or whitish. Leaves drying reddish brown or olive-green. Young twigs whitish grey to brown |
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Sepals fused for c. 3/4 of their length, apex acute; disc hirsute. Infructescences 14–28 cm long. BOR (Sabah) |
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91b. |
Sepals fused for c. 1/2 of their length, apex obtuse to rounded; disc sparsely pilose. Infructescences 10–75 cm long |
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Fruiting pedicels 0–1(–3) mm long. Fruits ovate to ellipsoid or lenticular, often apiculate, with a terminal to subterminal style. Ovaries hairy. Inflorescences and infructescences robust. Leaves usually drying yellowish brown. SUM, MLY, BOR, THA |
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92b. |
Fruiting pedicels 1–14 mm long. Fruits obliquely ovate (mango-shaped), with a lateral to subterminal style. Ovaries glabrous, very rarely hairy. Inflorescences and infructescences rather slender. Leaves drying olive-green. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
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Infructescences 18–45 cm long. Style lateral to subterminal, rarely terminal; calyx lobed for no more than 1/3 of its length, margin entire. Indumentum ferrugineous to ochraceous, rarely absent. Leaves drying reddish brown, tertiary veins usually perpendicular (at right angle to the midvein). Young twigs first dark brown, then light grey to whitish, or whitish from the beginning. Mostly rheophytic. BOR, PHI (Palawan), SUL |
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93b. |
Infructescences 6–20 cm long. Style terminal, rarely subterminal; calyx usually lobed for more than 1/3 of its length, margin often glandular-fimbriate. Indumentum absent or whitish, never ferrugineous. Leaves drying olive-green, tertiary veins oblique in relation to the midvein. Young twigs brown |
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Plant rheophytic, leaves 4–10 times as long as wide. SUM, MLY, BOR, THA |
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94b. |
Plant not rheophytic, leaves (2–)2.5–3.5(–6.6) times as long as wide.SUM, MLY,BOR, JAW,PHI, SUL, LSI, MOL, THA |
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Indumentum, especially of young parts and inflorescences, ferrugineous to ochraceous |
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95b. |
Indumentum whitish or absent |
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Fruits 6–20 by 4–8 mm, with a lateral to subterminal style. Leaves: petioles 1–5 mm wide. Infructescences (4–)7–65 cm long. Leaves (5–)9–30(–55) cm long, drying reddish brown to olive-green |
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96b. |
Fruits 3–5 by 3.5–4 mm, with a terminal to slightly sublateral style. Leaves: petioles 0.7–1.2 mm wide. Infructescences 3–7 cm long. Leaves (2–)4–10(–16) cm long, drying olive-green |
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Indumentum pale yellow to ochraceous. Ovaries globose to lenticular, spreading-hirsute, with a thin, conspicuous style. Fruits 6–8 by 4–6 mm, ellipsoid to lenticular, with a lateral style, often conspicuously beaked; fruiting pedicels 1–3 mm long. Infructescences (4–)8–24 cm long. Leaves 5–13(–20) cm long, drying greyish, reddish or olive-green. Stipules 3–7 by 0.5–1.2 mm, not falcate, coriaceous or strongly veined. Leaves: petioles 1–2 mm wide. LSI, MOL, SUL |
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97b. |
Indumentum ferrugineous to ochraceous. Ovaries ovoid, slightly falcate, appressed-pubescent, with an inconspicuous style. Fruits (7–)9–20 by 5–9 mm, obliquely ovate (mango-shaped), with a lateral to subterminal style; fruiting pedicels 0–23 mm long. Infructescences 7–65 cm long. Leaves (8–)15–60 cm long, drying reddish brown. Stipules (3–)6–35 by 1–10 mm, often falcate, coriaceous, strongly veined. Leaves: petioles (1–)2–5 mm wide |
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Plant rheophytic, leaves more than 6 times as long as wide. Leaves: petioles 1–3 mm wide. Inflorescences 3–5(–20) cm long, infructescences 7–20 cm long. Fruits 7–9 by 3–6 mm. BOR |
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98b. |
Plant not rheophytic, leaves less than 6 times as long as wide. Leaves: petioles 2–5 mm wide. Inflorescences (5–)10–30 cm long, infructescences 10–30 (–65) cm long. Fruits (7–)9–20 by 5–9 mm. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, THA |
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Sepals free, pubescent outside, c. 0.7 mm long. Fruits pilose. Inflorescences consisting of (1–)10–20 branches. Leaves pubescent at least along the major veins abaxially. Common. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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99b. |
Sepals fused for 1/2 of their length or more (splitting irregularly in fruit), glabrous outside, 1–1.5 mm long. Fruits glabrous. Inflorescences simple or once-branched, more rarely consisting of up to 6 branches. Leaves glabrous or nearly so. Rare. PHI |
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Fruits dorsiventrally compressed, 7–8 by 4–5 mm, pilose; disc, ovaries and abaxial leaf surfaces (at least along the veins) densely hirsute. MLY,THA |
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100b. |
Fruits laterally compressed or terete, 3–18 by 2.5–11 mm, glabrous or hairy; disc, ovaries and leaves glabrous or hairy |
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Fruits 8–18 by 6–11 mm, laterally compressed. Inflorescences 8–35 cm long, infructescences 10–75 cm long, simple. Leaves and inflorescence axes glabrous |
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101b. |
Fruits 3–8 by 2.5–6 mm, laterally compressed or terete. Inflorescences 2–10 cm long, infructescences 3–20 cm long, simple or branched. Leaves and inflorescence axes glabrous or hairy |
|
Fruiting pedicels 0–1(–3) mm long. Fruits ovate to ellipsoid or lenticular, often apiculate, with a terminal to subterminal style. Ovaries hairy. Leaves usually drying yellowish brown. SUM, MLY, BOR, THA |
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102b. |
Fruiting pedicels 1–14 mm long. Fruits obliquely ovate (mango-shaped), with a lateral to subterminal style. Ovaries glabrous, very rarely hairy. Leaves drying olive-green. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL |
|
Fruits terete, with a terminal style. Sepals often glandular-fimbriate. Leaves membranaceous to chartaceous, only larger leaves sometimes coriaceous |
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103b. |
Fruits laterally compressed, with a terminal to lateral style. Sepals never glandular-fimbriate. Leaves chartaceous to coriaceous |
|
Plant rheophytic, leaves 4–10 times as long as wide or narrower than 2 cm |
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104b. |
Plant not rheophytic, leaves (1.2–)2.5–3.5(–6.6) times as long as wide. SUM, MLY,BOR, JAW,PHI, SUL, LSI, MOL, THA |
|
Most leaves 5–11 by 1–2 cm. Leaf apex acuminate. SUM, MLY, BOR, THA |
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105b. |
Most leaves 3–6 by 0.4–1.2 cm. Leaf apex acute, sometimes with a rounded apiculum. THA |
|
Fruits 6–8 by 4–6 mm, with a lateral to subterminal, rarely terminal style. Leaves drying greyish, reddish or olive-green. Infructescences 4–24 cm long |
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106b. |
Fruits 3–5 by 2.5–4 mm, with a terminal to slightly subterminal style. Leaves drying olive-green. Infructescences 3–7 cm long |
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Stipules, Leaves: petioles and leaves glabrous. Young twigs light grey. Leaves chartaceous to coriaceous. SUM, MLY, THA |
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107b. |
Stipules, Leaves: petioles and abaxial leaf surfaces hirsute. Young twigs brown. Leaves membranaceous to chartaceous. LSI, MOL, SUL |
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Sepals free, pubescent outside, c. 0.7 mm long. Fruits pilose. Inflorescences consisting of (1–)10–20 branches. Leaves pubescent at least along the major veins abaxially. SUM, MLY, BOR, JAW, PHI, SUL, LSI, MOL, NWG, THA |
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108b. |
Sepals fused for 1/2 of their length or more (splits irregularly in fruiting stage), glabrous outside, 1–1.5 mm long. Fruits glabrous. Inflorescences simple or once-branched, more rarely consisting of up to 6 branches, sometimes fascicles of 2 inflorescences. Leaves glabrous or nearly so. PHI |