Malesian Euphorbiaceae Newsletter 3

 

 

 

 

 

Goto on this page:

Introduction

Important Literature

Progress

New Collaborators

Flora Malesiana Symposium

Changes in Malesian Euphorbiaceae Genus List

Genera and Species to be revised

Revised Genera

Additions and Changes in the Mailing List


Introduction

 

The third edition of the Malesian Euphorbiaceae Newsletter has been finished. It has been completely compiled again by myself. I like to emphasize that everybody is invited to send in short notes. I very much like to receive notes about progress, field trips, and other events. This way we can update each other about the state of affairs in the revision of the Malesian Euphorbiaceae.

The newsletter is becoming popular, the mailing list is still increasing. Also the number of participants is still increasing, only a few genera are left which are not yet reserved by a researcher for a revision.

In this edition of the newsletter I like to draw your attention to new literature, progress reports, and the Flora Malesiana Symposium

In the next issue special attention will be given to the Expert Center for Taxonomic Identification (ETI).

 

Important Literature

 

Recently, the first part of the proceedings of the International Conference on the Systematics of the Euphorbiaceae was published. The conference was held in St. Louis in August 1989. In this first part, of these very belated proceedings, Grady Webster presents a classification and synopsis of the Euphorbiaceae, containing keys to and descriptions of the infrafamily levels and the genera. Per taxon a full account of its synonymy is presented. The account does not as much provide a new critical delimitation within the Euphorbiaceae, but is more an overview of the presently (more or less) accepted genera. The family view is broad, most small split families have been included: Stilaginaceae (Antidesma), Pandaceae (Galearia, Microdesmis), and Bischofiaceae (Bischofia). Not included are the small families Buxaceae and Daphniphyllaceae. This concept of the Euphorbiaceae equals the one used for the genera list of Malesian Euphorbiaceae (see below). The genera list includes the Stilaginaceae, Pandaceae, and Bischofiaceae and excludes the Buxaceae and Daphniphyllaceae.

Webster's treatment incorporates a natural system, therefore, the keys follow the classification. For this reason, the keys may be important to identify Malesian specimens up to the genus level. The artificial key published by Airy Shaw (e.g., the issues on Bornean and New Guinean Euphorbiaceae) is difficult to use. Webster's key, with a completely different structure, might therefore be helpful. However, with both keys complete material (male and female flowers and fruits) is often still needed.

Other papers presented during the conference will be published in future issues of the Annals of the Missouri Botanical Garden.

Webster, G.L. 1994. Systematics of the Euphorbiaceae: Introduction. Ann. Missouri Bot. Gard. 81: 1,2.

Webster, G.L. 1994. Classification of the Euphorbiaceae. Ann. Missouri Bot. Gard. 81: 3-32.

Webster, G.L. 1994. Synopsis of the genera and suprageneric taxa of Euphorbiaceae. Ann. Missouri Bot. Gard. 81: 33--144.

 

Progress

 

Two small genera have been completed in the mean time, Chaetocarpus and Neoscortechinia. Especially the genus Neoscortechinia was difficult, the differences between the species are very small. Six species have been recognized, most of them are wide-spread in W Malesia. One variety will be raised to specific rank, this species occurs locally in NE Borneo. Another very wide-spread species will be split up into two, a W Malesian species and an E Malesian species (New Guinea and the Solomon Islands). The revision, which includes a phylogenetic analysis with Cheilosa montana as outgroup, will probably be published in the next issue of Blumea. Therefore, no names are presented here, otherwise the new combinations will have been made in this obscure 'journal'.

The genus Chaetocarpus, a pantropical genus, is only represented with one species in Malesia, Ch. castanocarpus (Roxb.) Thwaites (India to W Malesia). Two more species can be found in Sri Lanka (Ceylon). One of them, Ch. pubescens (Thwaites) Hook.f., resembles a specimen collected in Irian Jaya (W New Guinea). The latter specimen was sterile except for some fruit remnants in an attached paper bag. No taxonomic status has been given to this specimen. (Unfortunately, Irian Jaya is still a major undercollected area, but presently, two projects have started in the Bird's head of New Guinea, which might reveal more and better specimens.) The revision will probably be published in the Indian journal Rheedea. Ch. castanocarpus, although vegetatively not very attractive, has very special, beautiful fruits (stinging hairs) and flowers (stamens united in an androphore; stigma papillae dendritic).

Two weeks ago I received a letter from Prof. Nguyen Nghia Thin, that he has finished the revision of the genera Lasiococca and Strophioblachia. As soon as I have received his manuscripts extra attention will be given to his results. Hopefully, this might already be in the next issue of the newsletter.

 

New Collaborators

 

In the former newsletter the grants of the European Community were already mentioned. Two postdoctoral fellowships within the 'Botanical Diversity of the Indo-Pacific region Human Capital and Mobility Programme' have been assigned to the revision of Euphorbiaceae. Mrs. Petra Hoffmann will start with the genus Antidesma in Kew, England, for the next two years and Stefan Dressler will work on the genera Bridelia and Cleistanthus in Leiden, The Netherlands. They are both from the Humboldt University in Berlin and they will start their new jobs on September, 1st. Petra Hoffmann will, after having finished Antidesma, finish the revision of Actephila.

Two more M.Sc. students in Bogor, Indonesia, have started a revision of Euphorbiaceae genera. Mrs. Musayyinah and Mrs. Endah Woro Wiryani will work on the genera Koilodepas and Codiaeum respectively. Their supervision, just like with the other three M.Sc. students, will be in the experienced hands of Dedy Darnaedi, Johanis Mogea, Mien Rifai, Rugayah, and Elisabeth Widjaja.

In Leiden we have also started the application procedure for a Ph.D. fellowship within the 'Pathways from Asia to New Guinea' programme of the Dutch Science Foundation (NWO). This Ph.D. fellowship will include the revision of Baccaurea. After the revision a phylogenetic analysis will follow and the resulting data will be combined with those of the two other Ph.D. students in the programme working on insects. Finally, a historical biogeographic analysis will have to show, which geographical patterns are involved in distributions from SE Asia to New Guinea.

 

Flora Malesiana Symposium

 

The third Flora Malesiana Symposium will held from 10-14 July in Kew, England. Last week the second and final circular has been published, which includes the registration forms and the abstract forms. Copies can be obtained from the secretary of organising committee, Mr. B.J. Blewett (Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, United Kingdom).

The circular also contains information for those seeking funds for their participation. The circular urges everybody to apply for grants in their own country, but three British organisations might also be helpful in providing special grants. The Kew herbarium can assist in writing invitation letters.

Programme:

Registration will be on Sunday, 9 July (followed by a dinner at St. Mary's University College), which will be continued on Monday morning.

On Monday the opening will be followed by a lecture of a/the keynote speaker and the current state of Flora Malesiana. In the afternoon there will be workshops and contributed papers.

Tuesday will start with phytogeography and cladistic studies, followed again in the afternoon by workshops and contributed papers.

On Wednesday local flora projects will present themselves and a poster session will be held. At the end of the afternoon there will be short excursions and the Flora Malesiana Board Meeting.

Monographs and biological papers will be presented on Thursday morning, while in the afternoon there will be a Computer applications workshop and contributed papers. The special Symposium Dinner, with an after dinner speech by Dr. Whitmore, will be in the evening.

Finally, on Friday, papers on ethnobotany, conservation, and ecology will be followed by summaries of the workshops and the closure of the symposium.

For those interested, the week after the Flora Malesiana Symposium, the Holttum Memorial Pteridophyte Symposium will be organised in Kew and for those staying over from the FM III a special excursion is organised on Sunday.

 

Papers and Posters

Everybody who likes to contribute a paper or a poster (display boards of 2 by 1 m are available) should send an outline for consideration to the Scientific Programme coordinator, Mr. M.J.E. Coode (address see above). A special committee 'will select and advise authors whether their presentations have been designated papers or posters'.

 

Herbarium visit

Those people who like to visit the Kew herbarium should, at least one month in advance, contact Mr. B.J. Blewett (address see above), and those who like to visit the Natural History Museum (BM) should contact the Keeper, Dep. of Botany, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. Many visitors are expected and special arrangements have to be made to facilitate all wishes as much as possible.

 

Workshop

I like to draw your special attention to the workshops on large and complex families. I like to organise a Euphorbiaceae workshop on Monday or Tuesday afternoon. The workshops will take 2 hours, from 14.00--16.00 h. If possible, while further suggestions are welcome, I like to cover 5 topics during the workshop:

- Delimitation of infrafamily taxa, genera and species

- Characters

- Geographical patterns

- Local flora projects

- Computer aid

The last two topics will also be covered in special sessions or workshops, but it will be worthwhile to discuss our participation in local flora projects (Tree Flora of Sabah and Sarawak, Flora of Brunei, Flora of the Philippines, Flora of Thailand, etc.) and our use of special software (DELTA, ETI, etc.). I am planning to organise a demonstration of the ETI (Expert Center for Taxonomic Identification; see also the next newsletter) during the symposium and of our own database project LBASE, while there might be a special DELTA workshop. Especially, the DELTA and ETI software (free obtainable) will be very handy in subtracting data for local floras, if all of these floras more or less contain the same format.

The first two topics might be a bit outdated after the publication of the Missouri proceedings. However, (hopefully) many researchers will join the symposium who were absent in St. Louis and we might discuss the region specific problems, e.g. the delimitation between Cheilosa and Neoscortechinia, between Baccaurea and Aporosa, etc. Also, a discussion of the use of special character sets (e.g., stigma structures, DNA sequences) might be an important topic.

A discussion of biogeographical patterns might help to interpret variation, which will be useful especially for those not familiar with the Malesian region.

I would like to see different chairpersons for the various topics. And I like to invite you to join the symposium and especially the workshop and to volunteer yourself as a possible chairperson. Perhaps I can already stimulate potential candidates:

Delimitation: Grady Webster?

Special characters: Max van Balgooy, Kochummen, Anne Schot, Wurdack?

Geographical patterns: Max van Balgooy, Tim Whitmore, myself?

Local flora projects: Lolita Bulalaco, Kochummen, Enki Soepadmo?

Computer aid: Jeremy Bruhl, myself?

 

Registration

Join the symposium and register before 28th February 1995, otherwise you are charged extra. The symposium costs are £ 90.00 for full members (£ 40 for students, and £ 50 for accompanying persons), a stay in the St. Mary's University College ranges between £ 33.70 and £ 38.50 per night (including dinner and breakfast), the symposium dinner is £ 30 per person and the Wednesday excursion is £ 15 or £ 20 per person.

 

Changes in Malesian Euphorbiaceae Genus list

 

Due to Grady Webster's overview of the genera of Euphorbiaceae several changes have to be made in the list with the Malesian genera. These changes are made in the list below and typed in bold.

The genus Chorisandrachne has been added to Leptopus.

The genus Coccoceras has been added (split of from Mallotus, delimitation uncertain).

The genus Deutzianthus (including the genus Loerzingia) has been added.

Homalanthus has become Omalanthus (already indicated by Esser).

Octospermum has been added.

Plukenetia is added with the genus Pterococcus in its synonymy. Armbruster and Gillespie have likewise been moved as collaberators for Pterococcus.

The Malagasian genus Securinega has been excluded from the list; it has often been confused with Flueggea..

The genus Symphyllea has been included in the list, but it might be a synonym of Epiprinus..

The genus Synostemon has been removed as synonym of Sauropus.

 

Genera and Species to be revised

 

The following list shows the Malesian Euphorbiaceae genera in alphabetical order, the number of species estimated world-wide, estimated within Malesia, presence world-wide, and the contributor. This list is the updated version from the second newsletter

Genus

World

Malesia

Presence

Contributor

Acalypha

430

25

Pantropical

Levin

Actephila

35

6

Indomalesia to Australia

Hoffmann

Agrostistachys

9

5

India to W Malesia

Nais

Alchornea  

70

3

Tropical

Banka & Bulalacao

Aleurites

6

1

W Pacific

Bulalacao

Alphandia

3

1

New Guinea, W Pacific

Leiden students

Annesijoa

1

1

New Guina

Leiden students

Antidesma

160

60

Old world tropics and warm areas

Hoffmann

Aporosa

75

60

Indomalesia to Solomons

Schot

Ashtonia

2

2

Malay Penin., Borneo

Nais

Austrobuxus

17

1

Malesia to Fiji

Levin

Baccaurea

80

50

Indomalesia to W Pacific

Leiden Ph.D.

Baliospermum

6

1

India to central Malesia

Leiden students

Bischofia

1

1

Indomalesia

Leiden students

Blachia

12

1

India to central Malesia

 

Blumeodendron

6

6

Andamans, Malesia

Perumal

Borneodendron

1

N Borneo

Nais

Botryophora

0

1

SE Asia, W Malesia

Perumal

Breynia  

25

10

China to Australia and New Caledonia

Bruhl, Nais

Bridelia  

60

15

Old world tropics

Dressler

Cephalomappa

5

5

S China, Malesia

Widuri

Chondrostylis  

2

1

SE Asia, W Malesia

 

Choriceras  

1

1

S New Guinea, Australia

Forster

Chrozophora

12

1

Mediterranean, trop. Africa to India

Leiden students

Cladogynos  

1

1

SE Asia, Malesia

 

Claoxylon

 80

40

Old world tropics

 

Cleidion

25

3

Pantropical

Bulalacao

Cleistanthus

130

40

Old world tropics

Dressler

Clonostylis

0

1

Synonym of Spathiostemon

 

Cnesmone

10

1

Assam to W Malesia

Armbruster, Gillespie

Coccoceras 

4

4

Burma to Borneo

 

Codiaeum

6

10

Malesia to Pacific

Wiryani

Croton

750

35

Tropical and warm areas 

Ahumada

Dalechampia  

110

1

Warm areas, esp. America

Armbruster

Deutzianthus

2

1

Indochina and Sumatra

 

Dicoelia

3

1

W Malesia

Webster

Dimorphocalyx

12

5

Indomalesia to Australia

 

Doroxylon

1

1

Malesia

Leiden students

Drypetes

200

50

E Asia, S Africa

Levin

Elateriospermum

1

1

 S Thailand, Malay Pen.

Kochummen

Endospermum

13

10

SE Asia to Fiji

Kochummen

Epiprinus

6

1

Assam to W Malesia

 

Erismanthus

2

2

SE Asia to W Malesia

Webster

Erythrococca

30

1

Trop. & S Africa

 

Euphorbia

1600

30

Cosmopolitan

Vajravelu

Excoecaria

40

5

Old world tropics

Djarwaningsih

Fahrenheitia

4

1

India to central Malesia

Kochummen

Flueggea

13

2

Tropical

 

Fontainea

2

1

New Guinea, Australia, New Caledonia

Forster

Galearia

6

5

SE Asia to Solomons

Metilistina Sasinggla

Glochidion

300

150

Tropical

 

Hevea

9

1

Amazonia

Leiden students

Homonoia

2

1

SE Asia, Malesia

Leiden students

Hura

2

1

Trop. America

 

Hymenocardia

5

Africa, SE Asia to Sumatra

 

Jatropha

170

5

Tropical and warm areas, N America

Fernandez Casas

Kairothamnus

1

1

New Guinea

Levin

Koilodepas

10

5

India to Malesia

Ms. Muzayyinah

Lasiococca

3

1

E Himalayas to Malay Pen.

Nguyen

Leptopus

20

1

W Himalayas to Australia

Bruhl

Macaranga

240

125

Old world tropics

Whitmore

Mallotus

140

50

Old world tropics

Mohamad

Manihot

98

3

Tropical and warm America

Leiden students

Margaritaria

4

1

Tropical

 

Megistostigma

4

1

SE Asia, W Malesia

Armbruster, Gillespie

Melanolepis

2

1

SE Asia to Pacific

 

Micrococca

14

1

Old world tropics

 

Microdesmis

10

2

Trop. Africa, SE Asia, W Malesia

Leiden students

Moultonianthus

1

1

Sumatra, Borneo

Leiden students

Neoroeptera

1

1

NE Australia

 

Neotrewia

1

1

Malesia

Leiden students

Octospermum

1

1

New Guinea

 

Omalanthus

35

15

Trop. Asia to Australia

 

Omphalea

20

3

Tropical

Gillespie

Ostodes

4

1

E Himalayas to Borneo

 

Pachystylidium

1

1

India to Central Malesia

Armbruster, Gillespie

Pedilanthes

14

1

N to tropical America

Forster

Petalostigma

7

1

New Guinea, Australia

Forster

Phyllanthus

600

80

Tropical and warm areas

Mitra, Webster

Pimelodendron

8

4

Malesia

Kochummen

Plukenetia

15

1

Africa to Malesia

Armbruster, Gillespie

Ptychopyxis

13

10

Thailand to W Malesia, E New Guinea 

Murdoyuwono

Reutealis

1

1

Philippines

Leiden students

Richeriella

2

1

SE China to central Malesia

Leiden students

Ricinus

1

1

Europe to Africa and Middle East

Leiden students

Sapium

100

3

Tropical and warm areas to Patagonia

Kochummen

Sauropus

40

5

SE Asia, Indomalesia

Bruhl

Sebastiana

100

2

Tropical and warm areas

 

Spathiostemon

3

1

Thailand, W Malesia, New Guinea

Leiden students

Stillingia

30

1

Tropical and warm areas

 

Strophioblachia

2

1

SE Asia to central Malesia

Nguyen

Sumbaviopsis

1

1

Assam to W Malesia

Leiden students

Suregada

40

3

Old world tropics

 

Syndyophyllum

1

1

Sumatra, Borneo, New Guinea

Leiden students

Synostemon

0

0

Old world tropics

Synonym of Sauropus

Tapoides

1

1

Borneo

Leiden students

Trewia

2

1

Himalayas to Hainan

Leiden students

Trigonopleura

1

1

W Malesia

van Welzen, Bulalacao, On

Trigonostemon

45

15

Indomalesia

 

Wetria

1

1

SE Asia to W Malesia, New Guinea

Leiden students

 

Revised genera

 

Genus

World-wide

Malesia

Distribution

Publication

Chaetocarpus

11

1

Pantropical

Rheedea, 1995?

Cheilosa

1

1

W Malesia

Blumea 38 (1993) 161-166

Neoscortechinia

6

6

Burma to Solomons

Blumea, 1994?

 

Additions and Changes in the mailing list

 

Chakrabarty, T.

Central National Herbarium, Botanical Survey of India, P.O. Botanic Garden, Howrah 711 103, India.

Darnaedi, Dedy

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor.

Dressler, Stefan

Museum für Naturkunde der Humboldt-Universität zu Berlin, Institut für Spezielle Botanik und Botanische Sammlungen Arboretum, Späthstrasse 80/81, 12437 Berlin, Germany. He will treat Cleistanthus and Bridelia in Leiden (Sept.).

Forster, P.

Queensland Herbarium (BRI), Department of Environment & Heritage, Meiers Road, Indooroopilly, Queensland 4068, Australia. He will treat: Choriceras, Fontainea, Pedilanthus, and Petalostigma

Hoffmann, Petra

Museum für Naturkunde der Humboldt-Universität zu Berlin, Institut für Spezielle Botanik und Botanische Sammlungen Arboretum, Späthstrasse 80/81, 12437 Berlin, Germany. She will treat Actephila and Antidesma (latter in Kew, Sept.).

Levin, G.A.

Change of address: Center for Biodiversity, Illinois Natural History Survey, 607 East Peabody Drive, Champaign, Illinois 61820, USA

Mogea, Dr. J.P.

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor.

Perumal, Balu

Department of Botany, School of Plant Sciences, University of Reading, Whiteknights PO Box 221, Reading RG6 2AS, England. He is treating Blumeodendron, Botryophora

Rifai, Dr. Mien A.

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor.

Rugayah, Mrs.

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor.

Vajravelu, Dr. R.

Arboretum, Department of Biology, University of Central Florida, Orlando, Florida 32816-0368, U.S.A. e-mail: fdrvajra@ucf1vm.cc.ucf.edu. Addition to address.

Widjaja, Elisabeth

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor.

Wiryani, Ms. Endah Woro

Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. She will revise the genus Codiaeum.

Wurdack, K.J.

Department of Biology, CB # 3280, Coker Hall, University of N Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, U.S.A. He is interested in DNA analyses.